Within the narrow walls of the Outdoor Activities Club room, the few but lively members carry on the camping spirit as they have fun with DIY club experiments. Exciting experiences such as these have brought Nadeshiko Kagamihara deep into the world of camping, making her want to once again go on a solo trip. However, other plans come first, as Ayano Toki and Rin Shima invite Nadeshiko to camp near Ooi River. Eager to go on their first journey together, the girls set off to forge precious memories that will stick with their hearts for years to come.

Himari Kino is a cheerful and outgoing girl who is able to make friends with anyone she wants. When she enrolls in high school and attends the freshman opening ceremony, Himari falls in love with Yori Asanagi, a reserved upperclassman, during Yori's band performance. After the show, Himari blurts out her love, but Yori is inexperienced when it comes to romance. Yori, who did not expect this outcome at all, looks for answers from her friends and music. She decides to respond to Himari's confession, and to her surprise, Yori discovers that Himari fell in love with her music and performance rather than Yori as a person. Despite the shock, Yori desires to make Himari truly fall in love with her. The duo spend time together after school on the rooftop, where Yori usually plays her guitar with hopes to change Himari's feelings for her.

For the past 13 years, the Nefarious Monster Army have appeared beneath their floating fortress every Sunday to advance their goal of conquering Earth. Luckily, the Dragon Keepers are here to save the day! Having defeated the majority of the monsters during their initial invasion, the heroes now routinely show up to clear away any remnants of resistance. Unbeknownst to the public, however, every fight beyond the first has been a deliberate show put on by both sides: the Dragon Keepers reap the fame and prestige from fighting imaginary enemies, while the surviving weakest monsters, altogether called Dusters, are allowed to live for another day. Sentouin D, a Duster tired of living this life of shame, leaves the fortress in an attempt to overturn his fate. With the unexpected help of the mysterious ranger Yumeko Suzukiri, he discovers the key to defeating the Dragon Keepers lies in their Divine Artifacts—ultimate weapons crucial to the rangers' transformation and unique powers. Teaming up with Suzukiri to infiltrate the garrison of Red Keeper Sousei Akabane, Sentouin D must defy all expectations to save his species from the cruel hands of humans.

Hotaru Hinase has lived nearly sixteen years without romance. She is satisfied alone as long as she can enjoy the little things in life with her beloved friends and family. In her mind, she is not meant to fall in love, nor does she understand the concept. While out with a friend, Hotaru witnesses the severe breakup of her schoolmate Hananoi, to whom she has never spoken. Afterward, she sees him sitting in the snow and holds her umbrella over him. To her surprise, this gesture results in him confessing his feelings for her at school the next day. Though Hotaru rejects him, Hananoi insists on letting her get to know him. He does whatever he can to woo her, from changing his hairstyle to finding her lost hairpin in the snow. Although she has no romantic feelings towards him, Hotaru believes that by spending more time with him, she will learn how to love. She agrees to date him and slowly begins to navigate what being a girlfriend entails, including how to reciprocate the kind gestures that Hananoi continues to perform for her.