Filled with an overwhelming sense of wonder for the world around her, Mari Tamaki has always dreamt of what lies beyond the reaches of the universe. However, despite harboring such large aspirations on the inside, her fear of the unknown and anxiety over her own possible limitations have always held her back from chasing them. But now, in her second year of high school, Mari is more determined than ever to not let any more of her youth go to waste. Still, her fear continues to prevent her from taking that ambitious step forward—that is, until she has a chance encounter with a girl who has grand dreams of her own. Spurred by her mother's disappearance, Shirase Kobuchizawa has been working hard to fund her trip to Antarctica. Despite facing doubt and ridicule from virtually everyone, Shirase is determined to embark on this expedition to search for her mother in a place further than the universe itself. Inspired by Shirase's resolve, Mari jumps at the chance to join her. Soon, their efforts attract the attention of the bubbly Hinata Miyake, who is eager to stand out, and Yuzuki Shiraishi, a polite girl from a high class background. Together, they set sail toward the frozen south. Sora yori mo Tooi Basho follows the captivating journey of four spirited girls, all in search of something great.

When Demon Queen Echidna begins her invasion of the human realm, the people turn to the hero Leo Demonheart to protect them. Blessed with insurmountable power, Leo easily repels Echidna's army, thereby saving the world. But instead of gratitude or admiration, Leo is met with disdain and scorn from his fellow humans, who fear his overwhelming strength and believe that it may eventually cause humanity's demise. Banished from the kingdom he once called home, Leo wanders aimlessly throughout the land until, one day, he hears rumors of Echidna's efforts to rebuild her army. Hoping that demonkind might accept him as an ally, he returns to the demon queen's castle and offers Echidna his help. Naturally, Echidna immediately rejects him. Still, not all hope is lost, as Leo manages to convince Echidna's four generals to let him secretly work under their supervision. Donning the identity of a masked dark knight named Onyx, Leo solves the demon army's problems one by one—gradually improving the quality of life around the castle. Above all else, however, Leo wants to accomplish one goal: to learn why Echidna started the war in the first place.

Two years after the battle against Eosmon, Davis Motomiya, Ken Ichijouji, Yolei Inoue, Cody Hida, T.K. Takaishi, and Kari Kamiya meet a boy named Rui Owada, who is found with a depowered Digivice and claims to be the first Digidestined ever. In addition, a Digimon named Ukkomon appears in the real world, who wants every person in the world to have their own Digimon partner.